The phrase "transversely what has changed painting" likely relates to shifts that affect painting in a non-linear or cross-cutting way rather than along a straightforward timeline or single dimension. The term "transversely" means acting or lying across something, or set crosswise, often at right angles to a principal axis. In the context of painting, a notable change described as occurring "transversely" may refer to how certain factors, such as ion migration in paint layers, affect painting surfaces laterally across colors or layers rather than only vertically through the layers. This has implications for the physical and visual changes observed in paintings over time.

Moreover, "transversely" might also metaphorically describe a broader cultural or conceptual change in painting, where artistic creativity now evolves in a transversal manner, blending different mediums, social themes, and techniques that cross traditional boundaries. Painting no longer merely represents but interacts dynamically with other art forms and social realities.

Thus, the key changes that have transversely affected painting combine physical/material transformations occurring across layers as well as transversal conceptual shifts in artistic practice and meaning.