The animal with the most chromosomes per cell among diploid, non-polyploid eukaryote species is the Atlas blue butterfly (Polyommatus/Plebicula atlanticus), which possesses 448–452 chromosomes (i.e. 224–26 pairs) per cell. However, the organism with the highest chromosome number recorded to date is the adders tongue fern Ophioglossum reticulatum, which has an estimated 1,440 chromosomes (or 720 pairs) per cell. The protist Oxytricha trifallax is thought to have around 15,600 chromosomes, albeit nanochromosomes. The white-bellied pangolin has 114 chromosomes, more than any mammal except the Bolivian bamboo rat, which has 118. Therefore, the Atlas blue butterfly has the most chromosomes among diploid, non-polyploid eukaryote species, while the adders tongue fern has the highest chromosome number recorded to date.